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Tim completed his PhD in molecular toxicologyÌýat Colorado State University, where he made use of flow cytometric techniques and developedÌýmultimethodÌýcomputational models to understand heterogenousÌýcellular states in response to differentÌýxenobiotics and in degenerativeÌýaging. Tim joined the Spencer Lab in June 2019 to delve deeper into single-cell quantifications and to investigate the variousÌýmechanisms and stressÌýresponsesÌýunderlying proliferativeÌýcellularÌýheterogeneity. Tim completed his post-doc in the Spencer Lab in August 2023 to take a position asÌýSenior Scientist for Calico Life Sciences in San Fransico, CA.